Complete Calendar


Calendar of Events

What's New/Course Changes

Problem set 8 is now posted.  It is due Dec. 13. 

The answers to problem set 6 are now posted on the Assignments page.  

 Here is the remaining discussion schedule.  

Date

Topic

Who

Dec. 8

Optimality

Ben, Dave

The syllabus for this year's class is here.  The assignments page is here

Complete Calendar

This schedule is approximate.  You should time your readings to coordinate with the material being discussed in class.

After each lecture is given, I will post my lecture notes here in pdf format.  Click on the lecture title to see the notes.
Lect. Date Topic Reading Assignments
1 Aug. 28 Kinds of evolutionary biology FW Chap. 1
2 Aug. 30 Hardy Weinberg, variance
3 Sept. 1 Covariance Rice pp.5-17
Sept. 4 Labor Day - no class
4 Sept. 6 Gamete frequencies Rice pp. 39-43
5 Sept. 8 Fitness FW Chap. 4
6 Sept. 11 Change in allele frequency

Natural selection in Maple

Maple demonstration

7 Sept. 13 Equilibria, Overdominance
Sept. 15 Discussion- How do you define and recognize adaptation?

Kenny Wray, Neil Aschliman

Gould and Lewontin. Proc. R. Soc. Lond. B Biol. Sci. 205:581-598.
Reeve and Sherman. Quart. Rev. Biol. 68:1-32.
Prob. set 1 due
8 Sept. 18 Adaptive landscapes, Frequency dependent selection

Paquin simulation

9 Sept. 20 Frequency dependence
10 Sept. 22 Mutation and natural selection  Rice 18-31 Prob. set 2 due
11 Sept. 25 Density dependence
12 Sept. 27 How to detect and measure natural selection
13 Sept. 29 Genetic drift Powerpoint by Kim van der Linde.

Houle lecture on same, plus appendix on matrix arithmetic.

Rice pp.73-93, 104-107
14 Oct. 2 Effective population size Rice Chap. 4
15 Oct. 4 Selection on two loci Rice pp. 44-72  Prob. set 3 due
Oct. 6 Discussion - Measuring selection Nielsen 2005

Schemske and Bradshaw 1999

16 Oct. 9 Epistasis and evolution
17 Oct. 11 Coadaptation, Mutation direct FW Chap. 3

 

18 Oct. 13 Mutation FW Chap. 5.  Individual Problem set (4) due
19 Oct. 16 More mutation 
20 Oct. 18 Neutral theory.  powerpoint  FW Chap. 7
21 Oct. 20 Mutation - indirect approaches.   Rice p.93-104

 

Oct. 23 Detecting departures from neutrality 

Akey figures

Rice  Chap. 5

FW Chap. 9

 

22 Oct. 25 Multiple forces Rice Chap. 6  

1st version of grant proposal

23 Oct. 27 Discussion Neutral theory, Amanda Kubes Smith and Keightley 2002 Nature 415:1022.

Fay et al.  2002 Nature 415: 1024

24 Oct. 30 Gene flow 
25 Nov. 1 Population structure
26 Nov. 3 Intro. to Quantitative genetics Rice, p. 189 -194 prob. set 5 due
Nov. 6 From one locus to many FW Chap. 18
27 Nov. 8 Discussion, Genetics of Adaptation, Katie, Amanda B.

 Sucena and Stern

Cresko et al. 

Comments on proposals of others
Nov. 10 Veteran's Day - No class
28 Nov. 13 Variances and assumptions  FW Chap. 19
29 Nov. 15 Estimating parameters Rice, 194-209
30 Nov. 17 Responses to selection.   Prob. set 6 due
31 Nov. 20 Measuring S.  Evolvability  Powerpoint. FW Chap. 20
32 Nov. 22 Selection on many traits.  Arnold et al. 2001
Nov. 24 Thanksgiving - no class
Nov. 27 Discussion. Contingency, Stephanie Schwinn Travisano et al.1995, Science 267:87.  Burch and Chao 2000. Nature 406:625. 
33 Nov. 29 Adaptive landscapes. Theories of adaptation. Process of adaptation. Prob. set 7 due
34 Dec. 1 Intro. to optimality. Parker and Maynard Smith, 1990 Nature 348:27.  
35 Dec. 4 Building an optimality model Rice Chap. 9
36 Dec. 6 Game theory, ESS. FW Chap. 23
Dec. 8 Discussion, Optimality Ben and Dave Herre 1987 Nature 329:627. Final paper due
Dec. 13 Final Individual problem set due

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